From left to right: a small village with about 150 people, the MediaWiki software and extensions, the crowd from Wikimania'14 and Wikipedia.
Here, I equate organelles to specialized sub-systems of the greater paramecium. In the case of Wikipedia, specialized sub-systems tend to take the form of software.
See also:
- Video: https://www.youtube.com/watch?v=-We4GZbH3Iw#t=34m05s
- Slides: https://commons.wikimedia.org/wiki/File:Wikipedia_is_socio-technical_--_Research_Showcase_(October,_2014).pdf